本作登场舞台「游戏业界」分为「神界」与「下界」两个不同的世界。神界是由绿色主机「雄伟的绿色大地～Leanbox」、黑色主机「厚重的黑色大地～Lastation」、白色主机「梦想的白色大地～Lowee」以及紫色主机「革新的紫色大地～Planeptune」等四块浮游大地所组成。在神界中四名女神有着激烈的派系斗争，由于女神的力量源自于自己所守护的大陆的居民（占有率）的数量，因此如何有效增加大陆的居民人数，将会是这场「主机战争」的关键所在。 Gamindustri: One of many parallel universes protected by four goddesses known as CPUs. Long ago, these four CPUs engaged in a war for supremacy known as the War of the Guardians. However, this war weakened their protective hold on the world inhabited by their people, and in turn it was flooded by monsters under the influence of an individual known as Arfoire.

During the War of the Guardians, one of the CPUs—Neptune—was defeated by the others and hurled down to the world’s surface from the